Phyllophaga er en meget artsrik slekt av biller som hører til gruppen oldenborrer (Melolonthinae) i den store gruppen skarabider (Scarabaeoidea).
Utseende Små til ganske store (12-35 millimeter), sylindriske, brunlige oldenborrer. Kroppen er forholdsvis tynnskallet og mangler skjell og påfallende hårkledning på oversiden.
Levevis Larvene lever i jorda som vanlig for oldenborrer. Noen av artene kan bli svært tallrike og gjøre skade i landbruket. De voksne billene gnager på mange slags planter.
Utbredelse Slekten er utbredt i Amerika, fra Argentina i sør til Canada i nord, inkludert øyene i Karibia. Rundt halvparten av artene lever i Nord-Amerika.
Systematisk inndeling
Ordenen biller, Coleoptera Underordenen Polyphaga Overfamilien skarabider, Scarabaeoidea Familien skarabider, Scarabaeidae Latreille, 1806 Underfamilien oldenborrer, Melolonthinae Samouelle, 1819 Stammen Rhizotrogini Burmeister, 1855 Slekten Phyllophaga Harris, 1827 Liste over artene
Phyllophaga abcea Saylor, 1940 Phyllophaga abdominalis (Moser, 1921) Phyllophaga abudantuni Chalumeau & Gruner, 1976 Phyllophaga acacoyahuana Morón & Blas, 2006 Phyllophaga acapulca Saylor, 1943 Phyllophaga acatlanensis Morón & Aragon-García, 2012 Phyllophaga aceitillar Woodruff, 2005 Phyllophaga acinosa (Arrow, 1920) Phyllophaga acunai Chapin, 1937 Phyllophaga adjuntas Saylor, 1940 Phyllophaga aegrotus Phyllophaga aemula (Horn, 1887) Phyllophaga aenea (Moser, 1921) Phyllophaga aeneotincta Chapin, 1932 Phyllophaga aequalis (LeConte, 1853) Phyllophaga aequatorialis (Moser, 1921) Phyllophaga aeruginosa (Burmeister, 1855) Phyllophaga affabilis (Horn, 1887) Phyllophaga afflicta (Blanchard, 1850) Phyllophaga aguadita Saylor, 1942 Phyllophaga ahlbrandti García-Vidál, 1988 Phyllophaga alayoi García-Vidál, 1978 Phyllophaga albina (Burmeister, 1855) Phyllophaga alcoa Woodruff, 2005 Phyllophaga aliada Sanderson, 1951 Phyllophaga aliciae Morón & Riva-Cervantes, 2005 Phyllophaga alquizara Chapin, 1932 Phyllophaga alvareztoroi Morón & Blas, 2006 Phyllophaga amazonica (Moser, 1918) Phyllophaga analis (Burmeister, 1855) Phyllophaga andersoni Smith & Paulsen, 2015 Phyllophaga androw Woodruff, 2005 Phyllophaga angulicollis (Bates, 1887) Phyllophaga angusta (Blanchard, 1850) Phyllophaga anolaminata (Moser, 1921) Phyllophaga anomaloides (Bates, 1887) Phyllophaga antennalis (Moser, 1921) Phyllophaga antiguae (Arrow, 1920) Phyllophaga anxia (LeConte, 1850) Phyllophaga apicalis (Blanchard, 1850) Phyllophaga apolinari Saylor, 1940 Phyllophaga apolinaria Saylor, 1942 Phyllophaga approxima Woodruff & Sanderson, 2005 Phyllophaga aragoni Morón, 2013 Phyllophaga arcta (Horn, 1887) Phyllophaga ardara Saylor, 1943 Phyllophaga arkansana (Schaeffer, 1906) Phyllophaga arribans Saylor, 1943 Phyllophaga arrowi (Saylor, 1935) Phyllophaga assmani Saylor, 1936 Phyllophaga atra (Moser, 1918) Phyllophaga atrata (Moser, 1918) Phyllophaga atratoides Morón, 2003 Phyllophaga audanti Wolcott, 1928 Phyllophaga aurea Luginbill & Painter, 1941 Phyllophaga austera (Erichson, 1847) Phyllophaga babis Saylor, 1943 Phyllophaga badbunnyi Cano, 2020 Phyllophaga bahiana (Saylor, 1935) Phyllophaga balia (Say, 1825) Phyllophaga balli Morón & Woodruff, 2008 Phyllophaga baneta Saylor, 1943 Phyllophaga baoruco Woodruff, 2005 Phyllophaga baracoana Chapin, 1932 Phyllophaga baraguensis Chapin, 1932 Phyllophaga barda (Horn, 1887) Phyllophaga baroni (Bates, 1890) Phyllophaga barrapotosiana Romero & Morón, 2017 Phyllophaga barroi Serrano & Morón, 2017 Phyllophaga barrosa Sanderson, 1951 Phyllophaga batillifer (Bates, 1887) Phyllophaga beameri Sanderson, 1958 Phyllophaga beckeri (Moser, 1921) Phyllophaga benexonana Morón, 2000 Phyllophaga bicavifrons Chapin, 1932 Phyllophaga bicolor (Moser, 1921) Phyllophaga bilobatata Saylor, 1939 Phyllophaga bilyi Chalumeau, 1989 Phyllophaga bimaculata García-Vidál, 1984 Phyllophaga bimammifrons Saylor, 1940 Phyllophaga bipartita (Horn, 1887) Phyllophaga blackwelderi Saylor, 1940 Phyllophaga blanchardi (Arrow, 1933) Phyllophaga blanda Sanderson, 1958 Phyllophaga bobevelynorum Schnepp, 2022 Phyllophaga bolacoides (Bates, 1887) Phyllophaga boliviensis (Blanchard, 1850) Phyllophaga bonfils Woodruff & Sanderson, 2005 Phyllophaga boruca Morón, 2003 Phyllophaga bowditchi Saylor, 1938 Phyllophaga brama Saylor, 1943 Phyllophaga bretti Smith & Paulsen, 2015 Phyllophaga brevicornis García-Vidál, 1988 Phyllophaga brevipilosa (Moser, 1918) Phyllophaga brownella Saylor, 1942 Phyllophaga bruneri Chapin, 1932 Phyllophaga bueta Saylor, 1940 Phyllophaga buapae Morón, 2013 Phyllophaga bucephala (Bates, 1887) Phyllophaga caanchaki Morón, 1998 Phyllophaga cahitana Morón, 2001 Phyllophaga calai Smith & Paulsen, 2015 Phyllophaga calceata (LeConte, 1856) Phyllophaga calderasa (Saylor, 1937) Phyllophaga callosiventris (Moser, 1921) Phyllophaga cambeforti Cartwright & Chalumeau, 1977 Phyllophaga campana Morón & Riley, 2005 Phyllophaga caneyensis García-Vidál, 1984 Phyllophaga canipolea Saylor, 1948 Phyllophaga canoa Sanderson, 1951 Phyllophaga canoana Morón, 2003 Phyllophaga capillata (Blanchard, 1850) Phyllophaga caraga Saylor, 1943 Phyllophaga cardini Chapin, 1932 Phyllophaga carnegie Woodruff, 2005 Phyllophaga cartaba Sanderson, 1951 Phyllophaga castaniella (Bates, 1887) Phyllophaga castineirasi García-Vidál, 1978 Phyllophaga catemacoana Morón, 2003 Phyllophaga caviceps (Moser, 1918) Phyllophaga cayennensis (Moser, 1918) Phyllophaga caymanensisSanderson, 1939 Phyllophaga cazahuata Morón, 2013 Phyllophaga cazieriana Saylor, 1938 Phyllophaga cececpana Morón, 2016 Phyllophaga centralis (Nonfried, 1894) Phyllophaga certanca Saylor, 1943 Phyllophaga chada Saylor, 1948 Phyllophaga chalumeaui García-Vidál, 1984 Phyllophaga chamacayoca Morón, 1991 Phyllophaga chamula Morón, 1999 Phyllophaga changuena Morón, 2003 Phyllophaga chiapas Saylor, 1943 Phyllophaga chiblacana Morón, 2003 Phyllophaga chimoxtila Morón, 2003 Phyllophaga chippewa Phyllophaga chiriquina (Bates, 1887) Phyllophaga chlaenobiana Saylor, 1936 Phyllophaga cholana Morón, 2003 Phyllophaga chortiana Morón, 2003 Phyllophaga cinnamomea (Blanchard, 1850) Phyllophaga citarae Morón & Neita-Moreno, 2014 Phyllophaga ciudadensis (Bates, 1887) Phyllophaga clavijeroi Morón, 2015 Phyllophaga clemens (Horn, 1887) Phyllophaga clypeata (Horn, 1887) Phyllophaga cneda Saylor, 1940 Phyllophaga coahuayana Morón, 2006 Phyllophaga cocleana Morón, 2013 Phyllophaga colima Saylor, 1943 Phyllophaga colimana (Moser, 1921) Phyllophaga collaris (Moser, 1921) Phyllophaga collaroides Morón, Lugo-Gracía & Aragon-García, 2015 Phyllophaga columbiana (Blanchard, 1850) Phyllophaga comaltepecana Morón, 2003 Phyllophaga cometes (Bates, 1887) Phyllophaga complexipennis García-Vidál, 1984 Phyllophaga conformis (Blanchard, 1850) Phyllophaga congrua (LeConte, 1856) Phyllophaga conicariana Morón, 2006 Phyllophaga contaminata Fall, 1932 Phyllophaga coronadis Saylor, 1941 Phyllophaga corrosa (LeConte, 1856) Phyllophaga costaricensis (Moser, 1918) Phyllophaga costura Sanderson, 1951 Phyllophaga crassa (Burmeister, 1855) Phyllophaga crassissima (Blanchard, 1850) Phyllophaga crena Saylor, 1941 Phyllophaga crenaticollis (Blanchard, 1850) Phyllophaga crenonycha Saylor, 1943 Phyllophaga crenulata (Frölich, 1792) Phyllophaga cribrosa (LeConte, 1853) Phyllophaga crinipennis (Bates, 1890) Phyllophaga crinitissima Wolcott, 1924 Phyllophaga cristobala Saylor, 1942 Phyllophaga cruces Morón & Riley, 2005 Phyllophaga cubana Chapin, 1932 Phyllophaga cupuliformis Langston, 1924 Phyllophaga curialis Reinhard, 1939 Phyllophaga cuyabana (Moser, 1918) Phyllophaga cylindrica (Burmeister, 1855) Phyllophaga darlingtoni Sanderson, 1940 Phyllophaga dasypoda (Bates, 1887) Phyllophaga davidsoni Woodruff, 2005 Phyllophaga davisi Langston, 1927 Phyllophaga debilis (LeConte, 1856) Phyllophaga delata (Horn, 1887) Phyllophaga deleri Smith & Paulsen, 2015 Phyllophaga delphinicauda Androw, 2016 Phyllophaga delplanquei Chalumeau & Gruner, 1976 Phyllophaga densata (Moser, 1918) Phyllophaga densepunctata (Moser, 1918) Phyllophaga dentex (Bates, 1887) Phyllophaga denticulata (Blanchard, 1850) Phyllophaga dieteriana Deloya & Morón, 1998 Phyllophaga diffinis (Blanchard, 1850) Phyllophaga diminuta Evans, 2003 Phyllophaga disca Saylor, 1943 Phyllophaga discalis Chapin, 1935 Phyllophaga dispar (Burmeister, 1855) Phyllophaga dissimilis (Chevrolat, 1865) Phyllophaga divertens (Bates, 1887) Phyllophaga dominicensis Cartwright & Chalumeau, 1977 Phyllophaga drakii (Kirby, 1837) Phyllophaga dsaimana Morón, 2003 Phyllophaga dubitata García-Vidál, 1978 Phyllophaga duenas Saylor, 1941 Phyllophaga dugesiana Morón, Rivera & López, 2001 Phyllophaga dulcis (Bates, 1887) Phyllophaga durangoana (Moser, 1921) Phyllophaga durangosa Saylor, 1943 Phyllophaga ecostata (Horn, 1887) Phyllophaga ecuadorae Blackwelder, 1944 Phyllophaga edrileyi Morón, 2015 Phyllophaga eladio Woodruff, 2005 Phyllophaga elegans (Nonfried, 1891) Phyllophaga elenans Saylor, 1938 Phyllophaga elizoria (Saylor, 1937) Phyllophaga elongata (Linell, 1896) Phyllophaga emberae Morón, 2015 Phyllophaga emburyi Saylor, 1938 Phyllophaga eniba Saylor, 1943 Phyllophaga ephilida (Say, 1825) Phyllophaga epigaea (Wickham, 1903) Phyllophaga epulara Sanderson, 1958 Phyllophaga erota Saylor, 1942 Phyllophaga errans (LeConte, 1859) Phyllophaga espina Sanderson, 1951 Phyllophaga esquinada Sanderson, 1951 Phyllophaga etundutejiana Morón & Hernández-Cruz, 2013 Phyllophaga euryaspis (Bates, 1887) Phyllophaga exorata (Horn, 1887) Phyllophaga expansa Chapin, 1932 Phyllophaga explanicollis Chapin, 1935 Phyllophaga falcata Smith & Paulsen, 2015 Phyllophaga falta Phyllophaga farcta (LeConte, 1856) Phyllophaga favosa Saylor, 1941 Phyllophaga ferrugata (Moser, 1918) Phyllophaga ferruginea (Moser, 1918) Phyllophaga ferupilis Saylor, 1941 Phyllophaga fervida (Fabricius, 1775) Phyllophaga fesina Saylor, 1948 Phyllophaga fimbriata Cahpin, 1932 Phyllophaga fissilabris (Bates, 1887) Phyllophaga flavidopilosa (Moser, 1921) Phyllophaga floridana Robinson, 1938 Phyllophaga forbesi Glasgow, 1916 Phyllophaga forcipata (Burmeister, 1855) Phyllophaga forsteri (Burmeister, 1855) Phyllophaga fossoria Sanderson, 1951 Phyllophaga foveicollis (Bates, 1887) Phyllophaga foxii Davis, 1920 Phyllophaga fragilipennis (Blanchard, 1850) Phyllophaga franciscana García-Vidál, 1984 Phyllophaga fraterna Harris, 1841 Phyllophaga freeborni Saylor, 1937 Phyllophaga fucata (Horn, 1887) Phyllophaga fulvipennis (Blanchard, 1850) Phyllophaga fulviventris (Moser, 1918) Phyllophaga fusca (Frölich, 1792) Phyllophaga fuscipennis (Moser, 1918) Phyllophaga futilis (LeConte, 1850) Phyllophaga gaigei Sanderson, 1948 Phyllophaga galeanae Saylor, 1943 Phyllophaga garrota Sanderson, 1951 Phyllophaga gastonguzmani Morón, 2016 Phyllophaga gaumeri Bates, 1890 Phyllophaga gentryi (Saylor, 1936) Phyllophaga georgiana (Horn, 1885) Phyllophaga ghentata (Saylor, 1937) Phyllophaga ginigra Saylor, 1940 Phyllophaga glaberrima (Blanchard, 1850) Phyllophaga glabricula (LeConte, 1856) Phyllophaga gloriae Morón, 2013 Phyllophaga godwini Morón & Riley, 2005 Phyllophaga gracilis (Burmeister, 1855) Phyllophaga gracillima (Dalla Torre, 1912) Phyllophaga gramma Morón & Woodruff, 2014 Phyllophaga grancha Saylor, 1948 Phyllophaga gravidula (Moser, 1921) Phyllophaga grossepunctata (Moser, 1918) Phyllophaga guanacasteca Morón & Solis, 2000 Phyllophaga guanahacabibensis Serrano & Morón, 2017 Phyllophaga guanicana Smyth, 1917 Phyllophaga guapilana Saylor, 1935 Phyllophaga guapiles Saylor, 1941 Phyllophaga guapilesea Saylor, 1941 Phyllophaga guapoloides Morón & Solis, 2001 Phyllophaga guatemala Saylor, 1940 Phyllophaga guerrocans Saylor, 1938